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Starscream was betrayed by Megatron (used as bait) and so Starscream joined the Autobots in order to get back at him. He wasn't exactly a team player, though, as you might imaginge. He ended up going back to the Decepticons when Thrust convinced him that would be the best way to get back at Megatron, but that was mostly just to further Thrust's own agenda. When Unicron entered the picture, however, Starscream wanted the Autobots and Decepticons to team up. There's a bit more to the story but that's about the jist of it without spoiling anything major.
Sayonara.
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