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Leo. Without the head, the others will lose their direction due to the lack of a clear goal. Besides maintaining discipline and fairness, leaders must prioritize the interests of the team or their clan, they cannot easily let personal feelings sway them. When faced with the bigger picture or the need to punish members who have failed in their duties, even if he doesn't like the decisions he makes and feels pain, he has to do so if it's for the sake of the team.
But honestly, I don't like that “Without one of them means their doom.“ Who knows, they might find other ways to overcome it. In the IDW TMNT 20/20, Leo is no longer the leader, but he also become a mystic warrior, and still did his best to help his brothers.
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